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Salako, et al.

                V. paradoxa but also supports healthy and viable tree
                populations, as identified through could be found based
                on the thresholded vegetation indices. To determine the
                SZPI, the following steps were taken:
                (i)  The  predicted  SDM probability  raster  was
                   reclassified into two classes  by setting a suitability
                                            46
                   threshold at ≥60% probability: suitable (1) and not
                   suitable (0; less than the threshold). 19,22
                (ii)  Vegetation  indices  were  reclassified  based  on
                   optimal point (greenness peak) thresholds, derived
                   from a response-curve mode. This is the peak point
                   along the vegetation  index gradient  (e.g., NDVI,   Figure  4.  Models’  performances  of  random forest
                   GNDVI, modified NDWI [MNDWI]) and, therefore,    (RF), boosted regression trees (BRT), and artificial
                   is considered indicative of viable seed locations (1).   neural  network (ANN). Class ranges from 0 to 1,
                   Values below this threshold were classified as poor/  with 0 indicating very  poor model performance,
                   non-viable seed locations (0).                   0.5 not better than a random prediction, 0.7 – 0.8
                (iii) The SZPI maps were generated  by spatially    acceptable, and >0.9 outstanding.
                   combining the thresholded V. paradoxa suitability   Abbreviation: AUC: Area under the curve.
                   map with the thresholded vegetation  indices—
                   GNDVI, MNDWI, and NDVI. The resulting map
                   contained two classes: high-priority seed location   viable  and non-viable  seed location)  and threshold
                   (2) and poor/non-priority seed location (1).     ranges, respectively.  The predicted  optimal  threshold
                (iv) The SZPI maps were intersected with climate-based   for total  annual  precipitation and the wettest  months
                   SZMs to  identify  areas  of overlap  or  intersection   were at 1,000 mm and 250 mm, respectively, while that
                   with seed zones and associated land uses, such as   of the driest month should not be <0.4 mm of rainfall.
                   forest, parks, and reserves. It should be noted that   In terms of water  stress, it was predicted  to respond
                   the vegetation index thresholds used were assumed   positively as the value increased from 0.01, reaching a
                   to  be  species-specific  (V. paradoxa),  as  different   peak response at 0.2. Overall, V. paradoxa was predicted
                   species may respond differently to NDVI and other   to exhibit positive responses to most of the vegetation
                   vegetation  indices, potentially leading  to varied   indices  (NDVI, EVI,  soil-adjusted  vegetation  index
                   results.                                         [SAVI], and NDWI).
                                                                       Figure 5C presents the relative contributions of each
                3. Results                                          predictor  variable to the  distribution  of V. paradoxa as
                                                                    estimated by the BRT and RF models, along with the
                3.1. Models’ predictive performances                model consensus shown in Figure 5D. The ANN model
                Figure  4 presents the predictive  performances of the   was excluded from further analysis due to its poor
                three models’ algorithms across the evaluation metrics   performance. There was an agreement between the RF and
                (AUC, accuracy, and Kappa index). Both RF and BRT   BRT models regarding the importance of most variables.
                performed excellently  in all three metrics, with RF   Water-related variables, specifically annual precipitation,
                exhibiting a marginally higher performance. In contrast,   precipitation of the wettest month, and MNDWI, emerged
                ANN performed poorly in all the metrics; its AUC score   as the most important predictors. Among the vegetation
                indicates the prediction was not better than random.  indices, both models ranked the GNDVI as the most
                                                                    important vegetation phenology parameter. However,
                3.2. Estimation of the variable importance and      NDVI was ranked higher by the BRT model compared to
                responses of V. paradoxa to climate and vegetation   the RF model, while both SAVI and EVI were consistently
                indices/phenology parameters                        ranked as the least important by both algorithms.
                Figure  5A presents the partial  dependence  plots or
                response  curves  of  V.  paradoxa  distribution  along   3.3. Predicted spatial distribution of V. paradoxa
                the  environmental  gradients (climate  and  vegetation   and the mapping of seed-zone priority index
                indices), while Figure 5B and Table 4 demonstrate the   Figure 6A-D demonstrates the predicted distribution of
                optimal  threshold  (used  for  the  reclassification  into   V. paradoxa as modeled by RF, BRT, and ANN, and the



                Volume 22 Issue 4 (2025)                        96                           doi: 10.36922/AJWEP025210160
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